Xanthoria speciation

Tycker att jag hade en poäng med denna argumentation:
"The phylogenetic trees constructed from DNA sequence data resulted in paraphyly of X. calcicola, because X. aureola is nested inside the same clade, which was also the case in a previous study (Scherrer & Honegger 2003). There is a fundamental distinction between gene trees and species trees (Avise 2000, Rosenberg & Nordborg 2002). Paraphyly, or even polyphyly, in gene genealogies might be a temporary transitional state before finally reaching reciprocal monophyly, owing to incomplete lineage sorting of a genetically variable ancestral population (Avise 2000). Consequently, the genealogical paraphyly observed here cannot be taken as evidence that X. aureola and X. calcicola are conspecific, nor can it be interpreted to mean that colonization has proceeded from an inland rock-walls to maritime habitats. The morphological, chemical, and ecological differences along with the lack of shared haplotypes, on the other hand, indicate that X. aureola and X. calcicola are best treated as separate species."

Lindblom & Ekman (Myc. Res., 2005, sid 194)